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The most famous mass extinction was the disappearance of non-avian dinosaurs at the end of the Cretaceous, 66 million years ago (Mya), after ruling the Earth for 170 million years1,2,3. The best-supported extinction model is the impact of a large asteroid in the Yucatán Peninsula (Chicxulub, Mexico), which set off a global cataclysm and environmental upheaval4,5. Although evidence for an end-Cretaceous impact is indisputable6, most scientific debate has focused on whether the extinction was geologically abrupt or gradual7,8,9,10,11, whether it was caused by factors intrinsic to dinosaurs12,13,14,15 or by extrinsic physical drivers16,17,18 or both acting in concert19,20. If extrinsic events had a role, the question is whether this driver was terrestrial or extra-terrestrial21,22,23. It has proved harder to posit a convincing killing model that explains exactly how the dinosaurs, as well as many other groups24,25, vanished. And yet other groups of animals and plants survived through this singular, short-term crisis26,27. Could some groups have been teetering on the brink already? Furthermore, the extinctions coincide with a period of long-term environmental changes that resulted in remarkably high sea levels, cooling climates and the spread of new habitat types on land, as well as massive volcanic activity at the end of the Cretaceous16,19,20.
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